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In the wake of unknown Catastrophes in the heart of Terra, a mineral of unimaginable power has been discovered. With it, society's technology has made huge leaps and bounds, but the substance causes a deadly incurable disease, leading to a worldwide enslavement of the infected. As a rebellion is now on the rise, a pharmaceutical company races for a cure to save humanity.

In the year 2024, the world has collapsed. Grotesque monsters lurk amongst the ruins of Japan, while remaining people scrape together what they can to survive. Kiruko, an odd-job girl in Nakano, accepts a mysterious woman's dying wish to take a boy named Maru to a place called Heaven. Maru is convinced that there will be a boy there who looks exactly like him.

In a distant future, Tetsuro is a human boy who wants his body replaced with a robotic one. This is possible, but to do so he has to reach the Immortal Planet onboard the space train Galaxy Express 999. Maetel, a beautiful and mysterious blonde woman dressed in Russian style, joins him in the long journey through space. Every episode sees our heroes arriving in a new planet's space train station.

The future’s looking bleak. It’s a good thing there’s a new sheriff in town. Her name is Jo and she’s an enigma even to herself. Alongside her partners Sei, Amy, and Meg, Jo fights for the citizens who can’t defend themselves. Confronting corruption and twisted science, these gals are on fire—and they’re the best chance the wounded city’s got.