In a Polish shtetl, two young men who have grown up together betrothe their unborn children, ignoring the advice of a mysterious traveler not to pledge the lives of future generations. Soon after, one of them dies, and the wife of the other dies in childbirth. The children grow up in different towns, without ever knowing of the betrothal, but the power of the vow leads them to meet each other when they are marriageable. The young woman, Leah, is promised to another man, but Channon, the son of the father who died, is a practitioner of mysticism, and seeks to win his bride through sorcery.

A young woman poses as a man in a group of klezmer musicians in Poland.

Tevye is a dairyman in the Russian Ukraine early in the 20th century. He lives in a cabin outside Boyberik with his wife Goldie, his widowed daughter Tseytl, her two children, and his younger daughter, the unmarried Khave. Khave is being courted by Fedya, a Christian, the son of a local government official. Tevye warns Khave against romance and marriage outside her faith, but Fedya is persuasive too. What will Khave decide, how will Tevye react, and when the Tsar initiates a pogrom, will Tevye's friends come to his defense? Can the stubborn Tevye reconcile his heart and tradition?